I

»0

I

n

yv>

ENTOMOLOGICAL NEWS

QL'

VOLUME L, 1939

K

PHILIP P. CALVERT, PH. D., EDITOR

ASSOCIATE EDITORS : E. T. CRESSON, JR. R. G. SCHMIEDER, PH. D. V. S. L. PATE

ADVISORY COMMITTEE :

PHILIP LAURENT J. A. G. REHN

CHARLES LIEBECK JOHN C. LUTZ

J. CHESTER BRADLEY, PH. D. MAX KISLIUK, JR.

FRANK MORTON JONES, Sc. D. WM. W. CHAPMAN

PUBLISHED BY THE AMERICAN ENTOMOLOGICAL SOCIETY,

PHILADELPHIA, PA.:

THE ACADEMY OF NATURAL SCIENCES

LOGAN SQUARE

1939

The several numbers of the NEWS for 1939 were mailed at the Post Office at Philadelphia, Pa., as follows:

No. 1— January January 26, 1939

" 2— February February 9

" 3— March March 10

" 4— April April 13

" 5— May , May 10

" 6 — June June 16

" 7— July July 18

" 8— October October 6

" 9 — November Novembers

The date of mailing the December, 1939, number will be announced on the last page of the issue for January, 1940.

Subscriptions for 1939 are now due.

ENTOMOLOGICAL NEWS

JANUARY, 1939

s

((*

Vol. L

CONTENTS

Clark, Austin H. and Leila F.— Butterflies of a Wood Road at Suffolk,

Virginia 1

Hayes — A Bibliography of Keys for the Identification of Immature In- sects. Part 1. Diptera 5

Gloyd — A Synopsis of the Odonata of Alaska 1]

Blaisdell — A New Species of Coelus Eschscholtz. (Coleoptera: Tenebri-

onidae): 16

Bodenstein — A New Holopyga from the Western United States. (Hym-

enoptera: Chrysididae) . 19

Gyger— A Treatment for Crumpled Wings of Odonata Nymphs to Dis- close their Venation 21

Babiy — Apis griseocollis DeGeer — Bombus separatusCresson. (Hymen- optera: Bombidae)

Schwarz — A Substitute Name for Patera Schwarz (Hymenoptera: Mel- iponidae)

Robinson — A New Species of Aphodius from New Jersey (Coleoptera:

Scarabaeidae) 24

Current Entomological Literature 25

Frost — Occurrence of Aphodius scrofa Fabricius in Western Maine

(Coleo.: Scarabaeidae) 30

PHILADELPHIA, PA.

THE ACADEMY OF NATURAL SCIENCES, 1900 Race Street

Entered at the Philadelphia, Pa., Post Office aa Second Class Matter.

Acceptance for mailing at the special rate of postage prescribed for in Section 1.

Act of Octobrr J. 1917, mthorized January 15, 1921.

ENTOMOLOGICAL NEWS

published monthly, excepting August and September, by The American

Entomological Society. Philip P. Calvert, Ph.D., Editor; E. T. Cresson, Jr., R. G. Schmieder,

Ph.D., V. S. L. Pate, Associate Editors.

Advisory Committee: Philip Laurent, J. A. G. Rehn, Chas. Liebeck, J. Chester Bradley, PhD., Frank Morton Jones, Sc.D., John C. Lutz, Max Kisliuk, Jr., Wm. W. Chapman.

The subscription price per year of ten (10) numbers is as follows: United States and possessions, Central and South America $3.00, Canada

$3.15, Foreign $3.25, Single copies 35 cents.

ADVERTISING RATES: Full width of page. Payments in advance. One issue, 1 in., $ 1.20, 3 in., $ 3.00, 5 in., $ 5.00, 7 in., $ 7.00 Ten issues, " 8.00, " 25.00, " 45.00, " 60.00 SUBSCRIPTIONS. All remittances and communications regarding sub- scriptions, non-receipt of the "News" or of reprints, and requests for sample copies, should be addressed to

ENTOMOLOGICAL NEWS, 1900 Race Street, PHILADELPHIA, PA. MANUSCRIPTS AND ADVERTISEMENTS should be sent to the editor, Dr. P. P. Calvert, Zoological Laboratory, University of Pennsyl- vania, Philadelphia, Pa.

TO CONTRIBUTORS. The receipt of all papers will be acknowledged and if they are accepted they will be published as soon as possible. If not accepted, authors will be so advised and postage requested for return of manuscripts. Articles longer than six printed pages will be published in two or more installments, unless the author is willing to pay for the cost of a sufficient number of additional pages in any one issue to enable such an article to appear without division. Proof will be sent to authors.

Twenty-five extras (separates) of an author's contribution will be given free when they are requested; they will be "run of form," that is without removal of extraneous matter (parts of other articles at beginning and at end), folded, but unbound, uncut and without covers. Authors wishing more than 25 separates can obtain them, at the rates given at the bottom of this page, by ordering at the time of returning proof. When more than the twenty-five free separates are ordered, ALL the extras will be free of ex- traneous matter.

The making of blocks and printing all illustrations will be charged to authors. The editor will furnish cost of same when requested.

Stated Meetings of The American Entomological Society will be held at 8.00 P. M., in 1939, on the fourth Thursday of each month excepting June, July, August, November and December, and on the third Thursday of November and December.

Communications on observations made in the course of your studies are solicited; also exhibits of any specimens you consider of interest.

The printer of the "News" will furnish reprints of articles, without covers, over and above the twenty-five given free at the following rates: One or two pages, twenty-five copies, 35 cents; three or four pages, twenty-five copies, 70 cents; five to eight pages twenty-five copies, $1.40; nine to twelve pages, twenty five copies, $2.00; each half-tone plate, twenty-five copies. 30 cents; each plate of line cuts, twenty-five copies, 25 cents: greater numbers of copies will be at the corresponding multiples of these rates." Printed •oven for 50 copies, $4.00 or more, according to number of paces bound.

ENTOMOLOGICAL NEWS

VOL. L. JANUARY, 1939 No. 1

Butterflies of a Wood Road at Suffolk, Virginia.

By AUSTIN H. CLARK and LEILA F. CLARK, U. S. National Museum, Washington, D. C.

Seventy-three species of butterflies we have found along a half-mile stretch of abandoned woodland road on the north- western border of the Dismal Swamp parallel to a lumber rail- way on the outskirts of Suffolk, Virginia. Such a wealth of butterflies, including so many rarities, in a restricted and easily accessible locality is unusual and seems worth recording. In- deed, more species undoubtedly are to be found there, for our visits to the region have been few, and our time on each visit limited to a few hours. Furthermore, every visit has yielded species we had not seen before, and in the more or less im- mediate vicinity we have found nine additional species that might well occur there.

Perhaps the most interesting butterfly here is Enodia creola which is found in the cane (Arundinaria gigantea) along the road for a distance of about 150 feet. Here we once took twenty-one specimens in a couple of hours. Since then we have paid little attention to it, but have noted its constant pres- ence. Everywhere along the road, except in this special region, E. portlandia is abundant.

Inconspicuous and always keeping near the ground, feeding on the flowers of violets, Prunella or Elephantopns according to season, is the little skipper Amblyscirtcs Carolina. It is not very common, but you may see as many as a dozen in the course of a morning. Usually you will see three or four — sometimes none. Much more numerous and conspicuous is A. iextor, which is sometimes abundant. On two brief visits two days apart, just before the middle of June, Mr. and Mrs. E. L. Bell secured sixty and we secured fifty-eight, after which we passed them by.

t

2 ENTOMOLOGICAL NEWS fj^»-. '39

From time to time you notice the strangely inert Atrytonc dion in the grass, and occasionally the wary Poanes ychl or Atrytonc logan, the last named of large size. Incidentally it is the northern form of Atrytone dion that is found here, not the smaller, darker, southern form (alabamac) found further northward in the Dahl Swamp in Accomac County on the Eastern Shore.

Near the trees burdened with mistletoe on the southern edge of the road you sometimes notice the unsuspicious and stupid Atlides Jialcsus, and in the grass, if you look closely, you may find Nymphidia pinnila perched head downward with wings outspread.

Naturally, all of the butterflies we have found here do not occur at the same season, nor are they equally abundant in different years. Most of the more unusual ones are commonest in a wet season- — rain with intervals of sunshine.

In the course of our investigations we have covered every section of Nansemond, Norfolk and Princess Anne Counties. Most of this area, more or less extensively drained and inten- sively cultivated, is singularly devoid of butterflies, even of the commonest species. When butterflies do occur they are mainly the species of barren country and waste lands, or of weedy roadsides. In the height of the season man)' gardens yield no butterflies at all, while in others you find only Vanessa virgini- ensis, Euptoieta clandia, Picris rapae, Atalopedes campcstris, Hylephila phylaens and Panoquina ocola, with an occasional swallowtail, usually Papilio gloncns or P. troilus, and some- times Terms lisa and Phocbis cnbitlc. Only in localized and usually widely separated regions are the more interesting species found.

There are some other spots that are very rich, perhaps as rich as this woodland road, but we have not worked them so intensively. And still others are notable for the occurrence in numbers of a particular species.

As an example, along the main highway (route 10) about two miles west of Spring drove, Surry County, we have found Argynnis diaua more numerous than we have ever seen it else-

1, '39] ENTOMOLOGICAL NEWS

where. Mrs. Barnes, who very kindly gave us permission to look over her garden, told us that she had seen as many as twenty-five at one time about her butterfly-bush.

The wood road is reached as follows : Starting from the Hotel Elliott opposite the Post Office at Suffolk you go southeast on Main Street for two blocks, then turn left onto Washington Street. Following Washington Street for about a mile you come to a fork with an Esso station in it. Bearing to the right on the White Marsh road past the Esso station you come in less than a mile to a lumber railway that crosses the road. Just beyond this railway crossing, on the left, is a large lumber yard. Turn to the left at the company store and, securing per- mission from the "boss-man," park your car in the lumber yard. Going to the lumber railway, you will see that it goes down a short incline into the Dismal Swamp. At the bottom of the incline leave the railway, cross the dry ditch on its north side and, passing through the brush, you immediately find yourself on the wood road.

The butterflies for which we have records from along this road and the nearby lumber railway are the following :

SATYRIDAE: Neonympha gemma (Hiibner), N. eurytus (Fabricus), N. sosybius (Fabricius), Cercyonis dope alope (Fabricius), Enodia portlandia portlandia (Fabricius), E. creola (Skinner).

NYMPHALIDAE: Polygonia interrogation's (Fabricius), P. comma (Harris), Nymphalis antiopa creta (Verity), Vanessa atalanta (Linne), V. virginiensis (Drury), V. cardui (Linne), Precis coenia Hiibner, Basilarchia arthemis astyanax (Fabri- cius), B. ar chip pus (Cramer), Phyciodes tharos (Drury), Argynnis diana (Cramer), A. cybele (Fabricius), Euptoieta claudia (Cramer).

DANAIDAE: Danaus plexippus (Linne).

RIODINIDAE : Nymphidia pumila Boisduval and LeConte.

LYCAENIDAE: Lycaenopsis argiolus pseudargiolus (Bois- duval and LeConte), Everes comyntas (Godart), Atlidcs hale- sus (Cramer), Strymon m-album (Boisduval and LeConte), Strymon cecrops (Fabricius), Strymon melinus (Hiibner).

PIERIDAE: Pieris rapae (Linne), P. protodice Boisduval and LeConte, Euchloe genutia (Fabricius), Phocbis cnbitlc (Linne), Colias philodice philodicc Godart, C. p. citi-ylhcinc

4 ENTOMOLOGICAL NEWS [Ja^L, '39

Boisduval and LeConte, Terias nicippe (Cramer), T. lisa (Bois- duval and LeConte).

PAPILIONIDAE : Papilio philenor Linne, P. polyxcncs asterius Cramer, P. crcsphontes Cramer, P. glaucus Linne, P. troilus Linne, P. palamedes Drury, P. marcellus Cramer.

HESPERIIDAE : Epargyreus clarus (Cramer), Achalarus lyci- ades (Geyer), Thorybes bathyllus (Smith), T. pylades (Scud- der), T. confusis Bell, Pyrgus communis (Grote), Pholisora catullus (Fabricius), Erynnis icelns (Scudder and Burgess), E. brizo (Boisduval and LeConte), E. juvenalis (Fabricius), E. horatius (Scudder and Burgess), E. tcrentius (Scudder and Burgess), Ancyloxypha numitor (Fabricius), Hylephila ph\- laeus (Drury), Atalopedes campestris (Boisduval), Polites verna (W. H. Edwards), P. manataaqua (Harris), P. thcmi- stocles (Latreille), P. peckius (Kirby), Wallengrenia otho egeretnet (Scudder), Poanes zabulon (Boisduval and Le- Conte), P. yehl (Skinner), Airy tone aragos (Boisduval and LeConte), A. logan (W. H. Edwards), A. dion (W. H. Ed- wards), A. ruricola (Boisduval), Lerema accius (Smith), Am- blyscirtes textor (Hiibner), A. Carolina (Skinner) (and var. reversa Jones), Lerodea I'herminier (Latreille), Panoquina ocola (W. H. Edwards).

The butterflies of this region that we have not found along this woodland road are: Neonympha areolatus areolatus (Hiib- ner), found a few miles southwest; A^. a. septentrionalis Davis, a few miles south and southwest; Satyrodes eurydice (Linne), a few miles west; Dione vanillae incarnata Riley, a few miles northeast; Mitoura gryneus (Hiibner), a few miles northwest; Urbanus proteus (Linne), a few miles east; Pholisora hay- hurstii (W. H. Edwards), a couple of miles south; Erynnis martialis (Scudder), a few miles south; Poanes viator (W. H. Edwards), a few miles northwest; Atrytone dukesi Lindsey, gum swamp at North Landing; Lerodea, eufala (W. H. Ed- wards), within a mile, in open fields; Calpodes ethlius (Cramer), a pest on canna in Suffolk in 1937.

Some of these undoubtedly occur along the woodland road from time to time, but others live in specialized habitats from which they do not stray. Still other species have been recorded from this region, but have not been found by us.

Mr. Frank Morton Jones deserves the credit for first having

1, '39] ENTOMOLOGICAL NEWS

called attention to this region, he having collected along the lumber railway. Mr. Ernest L. Bell was the next visitor. On two days last June we had the pleasure of visiting the wood road in company with Professor and Mrs. Charles T. Brues and Miss Alice Brues, and Mr. and Mrs. E. L. Bell. Others who have collected in this locality are Dr. G. W. Rawson of Detroit, Mr. W. Herbert Wagner of Washington, and Mr. John Boyd of Southern Pines, N. C. All of these have been so very kind as to send us their records.

A Bibliography of Keys for the Identification of Immature Insects. Part I. Diptera.

By WM. P. HAYES, University of Illinois.

(Continued from Vol. XLIX, page 251.) SlMULIIDAE.

EDWARDS, F. W. 1920. On the British species of Simulium II. The early stages ; with corrections and additions to Part I. Bull. Ent. Res., 11:211-246. (Key to species of larvae and pupae, pp. 219-223.)

ID. 1934. The Simuliidae (Diptera) of Java and Sumatra. Archiv. Hydrobiol, Band 13, Suppl. Heft. 1, pp. 92-138. (Lar- val and pupal keys to species, pp. 101-103.)

EMERY, W. T. 1913. The morphology and biology of Simulium vittatum and its distribution in Kansas. Kansas Univ. Sci. BuL, 8:323-362. (Keys to larvae and pupae, pp. 350-351.)

JOHANNSEN, O. A. 1903. [Simuliidae of the United States.] N. Y. State Mus. Bui. 68, pp. 336-388. (Key to larvae and pupae, pp. 353-354.)

LUTZ, A. 1910. Segunda contribuicao para o conhecimento das especies brazileiras do genero Simulium. Mem. Inst. Os- waldo Cruz. 2:213-267. (Key to pupae of Brazilian species in Spanish and German, pp. 263-267.)

MALLOCH, J. R. 1914. American black-flies or buffalo-gnats. U. S. Dept. Agr., Bur. Ent. Tech. Bull. 26. (Table of larvae, p. 14, table of pupae, p. 15.)

PURI, I. M. 1925. On the life history and structure of the early stages of Simuliidae (Diptera). Parts I and II. Para- sltology, Vol. 17, Pt. I, pp. 295-334, Pt. II, pp. 335-369. (In part II, key to full grown British larvae and pupae, pp. 334- 337.)

6 ENTOMOLOGICAL NEWS [Jan., '39

TWINN, C. R. 1936. The black-flies of eastern Canada (Simuliidae, Diptera). Canad. Jour. Res. 14; Sec. D., pp. 97- 130 and pp. 131-150. (Key to pupae.)

CHIRONOMIDAE.

SPARCH, R. 1922. Bietrage zur Kenntnis der Chironomiden- metamorphose. I-IV. Part I. Kronologische Ubersicht der bisher veroffentlichten Litteratur iiber Chironomidenlarven und Chironomidenmetamorphose. Ent. Mcddcl. 14:32-109. (This has no keys but should be consulted by anyone interested in the life stages of this family,— a chronological history and good bibliography).

JOHANNSEN, O. A. 1905. Mayflies and midges of New York. N. Y. State Mus. Bull. 86: 76-331. (Keys to species of larvae and pupae, pp. 87-88.)

ID. 1937. Aquatic Diptera, Part III. Chironomidae : Sub- families Tanypodinae, Diamesinae, and Orthocladiinae. Cornell Agr. Ex p. Sta. Mem. 205, pp. 1-84, 18 pi.

KARNY, H. H. 1934. Biologic der Wasserinsekten. Fritz Wagner, Wien, 311 pp. (Key to genera and species of Tendi- pedidae-Chironomidae, pp. 208-213.)

KRUGER, F. 1938. Tanytarsus-studien I. Die subsectio Atanytarsus. Zugle;ch variationsstatistiche Untersuchungen zuni Problem der Artbildung bei Chironomiden. Arch. Hydro- bioL 33:208-256. (Key species of larvae of Cladotanytarsus, pp. 221-222.)

LENZ, F. 1921. Chironomidenpuppen und Larven. Bestim- mungstabellen. Deutsche Ent. Zeit., 1921, pp. 148-162. (Keys to larvae and pupae.)

MALLOCH, J. R. 1915. Chironomidae or midges of Illinois, etc. Bull. III. State Lab. Nat. Hist., 10:275-543. (Keys to larvae and pupae, pp. 364-366 and 401-410.)

ID. 1917. Preliminary classification of Diptera, etc. Part I. (Part II was never published.) Bull. ///. State Lab. Nat. Hist., 12: 161-409. (Keys to subfamily, genera and species of larvae and pupae, p. 286-290.)

THIENEMANN, A. 1908. Uber die Bestimmung der Chiro- nomidenlarven und Puppen. Zool. Anz. 33:753-756. (Key to genera of larvae and pupae, pp. 754-756.)

ID. 1914-1921. Vorarbeiten fur eine Monographic der Chironomiden-Metamorphose. Archiv. Hydrobiol. Suppl. Band II. 850 pp. This work, edited by Thienemann, contains the following papers by various authors which contain keys to Chironomidae . BAUSE, E. 1914. Die Metamorphose der Gattung Tanytarsus

1, '39] ENTOMOLOGICAL NEWS

und einiger verwanter Tenipediden-arten. pp. 1-128. (Lar- val key to species, pp. 42-46, pupal key to species, pp. 8S 53.) GRIPEKOVEN, H. 1914. Minierende Tendipediden. pp. 129-230.

(Various keys to larvae and pupae of these leaf -mining

Chironomids.) POTTHAST, A. 1915. Uber die Metamorphose der Ortho-

cladius-Gruppe. pp. 243-376. (Key to species of larvae and

pupae, pp. 252-257.) REITH, J. T. 1915. Die Metamorphose der Culicoiden (Cera-

topoginen). Pp. 377-442. (Various keys to larvae and

pupae.) THIENEMANN, A. 1921. Metamorphose der Tanypinen. Part

I, pp. 566-654; Part II, pp. 655-784. (Keys to larvae and

pupae, pp. 667-674; see also tables after p. 644.) ID. 1921. Die Metamorphose der Chironomidengattungen

Camptocladius, Dyscamptocladius und Phaenocladius mit

Bemerkungen uber die Artdifferenzierung bei den Chiro-

nomiden uberhaiipt. Pp. 809-850. (Various keys to larvae

and pupae of these genera.)

THIENEMANN, A. 1924. Uber die Chironomidengattung Lundstromia nebst einer Bestimmungstabelle fiir die Larven und Puppen der Sectio Tanytarsus. Zool. Anz. 58:331-345. (Key to genera and species of larvae and pupae, pp. 331-345.)

ID. 1928. Chironomiden-Metamorphosen I. Archiv. Hydro- biol. 19: 585-623. (Various keys to larvae and pupae of Cera- topogininae.)

ID. 1929. Chironomiden Metamorphosen II-Die Sectio Tanytarsus genuinus. Archiv. Hydrobiol. 20:93-123. (Vari- ous keys to larvae and pupae of this group.)

ID. 1934. Chironomiden-Metamorphosen III. Zur metamor- phose der Orthocladiariae. Dent. Ent. Zeit. for 1933, p. 1-38. (Various keys to larvae and pupae.)

ID. 1935. Chironomiden-Metamorphosen. X. Orthocladius- Dactylocladius (Diptera). Stettin. Ent. Ztg. 96:201-224.

ID. 1935. Chironomiden-Metamorphosen XII. Deutsche. Ent. Zeit. Jahrgang 1935 Heft, 1 and 2, p. 86-96. (Pupal key to sp. of Brill a, p. 94.)

ID. 1936. Chironomiden-Metamorphosen XI. Die Gattung Eukiefferiella. Stett. Ent. Zeit. 97: 43-65. (Larval key pp. 45- 47 and pupal key pp. 48-50.)

ID. 1936. Chironomiden-Metamorphosen XIII (Diptera). Die Gattung Dyscamptocladius Thien. Mitt, der Dent. Ent. Gesell. E. V. 7:49-54. (Key to species of larvae and pupae, pp. 50-52.)

8 ENTOMOLOGICAL NEWS [Jan., '39

ID. 1937. Chironomiden aus Lapland. III. Beschreibung neuer Metamorphosen, mit einer Bestimmungstabelle der bisher bekannten Metriocnemuslarven und-puppen. Stett. Ent. Zeit., 98:165-185. (Larval and pupal keys to species of Metrioc- nemus.)

THIENEMANN, A. and O. HARNISCH, 1932. Chironomiden Metamorphosen IV. Die Gattung Cricotopus. Zool. Am:. 99: 133-143. (Keys to a few genera included in the Cricotopus group. )

THIENEMANN, A. and K. MAYER. 1933. Chironomiden Metamorphosen VI. Zool. Anz., 103: 1-12. Larval and pupal keys to species of the Diamesa group, pp. 9-12.)

ZARVEL, J. 1928. Die Jugendstadien der Tribus Corynoneu- rariae. Archiv. Hydroblol. 19:651-665. (Keys to larvae and pupae of genera and species, pp. 662-665.)

ID. 1933. Larven und Puppen der Tanypodin von Sumatra und Java. Archiv. Hydrobiol, Suppl. Band 11, pp. 604-624.

ID. 1934. Tanytarsus larven und puppen aus Niederlandisch- Indien. Archiv. Hydrobiol., Suppl. Band 13. Heft 1. Pp. 139- 165. (Contains various keys to species of larvae and pupae.)

CERATOPOGONIDAE.

KARNY, H. H. 1934. Biologic der Wasserinsekten. Fritz Wagner, Wien, 311 pp. (Key to genera, pp. 207-208.)

KIEFFER, J. J. 1901. Zur Kenntniss der Certopogon-larven. Algm. Zeit. Ent., 6: 216-220. (Key to species, pp. 216-217.)

LENZ, F. 1934. Die Metamorphose der Heleidae. In : Die Fliegen der Palaerktischen Region by E. Lindner. Lief. 78: 95-128. (Key to genera of larvae and pupa, pp. 98-99; and various keys to species.)

MALLOCH, J. R. 1915. Chironomidae, or midges of Illinois. Bull III. State Nat. Hist. Surv., Vol 10, pp. 275-543. (See also Malloch, 1917, in general list.)

MAYER, K. 1934. Ceratopogoniden-Metamorphosen (C. in- ter mediae and C. vermiformes) der Deutschen Limnologischen Sundra-Expedition. Archiv. Hydrobiol., Suppl. Band 13, Heft 1, pp. 166-202. (Keys to eggs, larvae and pupae of the two groups cited in title, pp. 170-172 and 182-185.)

REITH, J. T. 1915. Die Metamorphose der Culicoiden (Cera- topogonien). Archiv. Hydrobiol., Suppl. Band 2, pp. 377-442. (Various keys to larvae and pupae.)

THIENEMANN, A. 1928. Chironomiden Metamorphosen I. Archiv. Hydrobiol, 19: 585-623. (Various keys to larvae and pupae.)

1, '39] ENTOMOLOGICAL NEWS

PSYCHODIDAE.

HALIDAY, A. H. 1856. Psychodidae. In: Walker, British Diptera, Vol. 3, pp, 253-263. (Key to genera of larvae, p. 25.)

HAZEMAN, L. 1907. A monograph of North American Psy- chodidae. Trans. Amcr. Ent. Soc., 33: 299-333. (Key to three European larval genera, p. 325.)

MIALL, L. C. and N. WALKER. 1895. The life-history of Pericoma canescens (Psychodidae). Trans. Ent. Soc. Land., for 1895, pp. 141-153. (A reprint of Haliday's key, p. 150.)

MUTTKOWSKI, R. A. 1915. New insect life histories I. Bull. Wis. Nat. Hist. Soc., 13: 109-122. (Key to North American larvae, pp. 111-112.)

See also: Johannsen, 1934, and Malloch, 1917, in general list.

DlXIDAE.

GOETGHEBUER, M. 1921. Les Dixidae de Belgique. Bull. Soc. Ent. Belg., 3: 183-188. (Key to larvae; paper not seen.)

SMITH, F. K. 1928. Larval characters of the genus Dixa. Jour. N. Y. Ent. Soc., 36: 263-284. (Key to species of larvae.)

See also: Johannsen, 1903 and 1934, in general list.

CULICIDAE.

BRAGA, J. M. 1931. Culicideos de Portugal. Aranjo and Sobrinko, Porto. (Keys to larvae of Culex, pp. 31-32; Aedes, pp. 52-53, and Anopheles, pp. 73-74.)

BUXTON, P. A. and G. H. E. HOPKINS. 1925. The early stages of Samoan Mosquitoes. Bull. Ent. Res., 15:295-301. (Key to larvae and pupae, p. 301.)

CHRISTOPHERS, S. R. 1933. Culicidae, tribe Anophelini. In: Fauna of British India. Diptera, Vol. 4. (Key to larvae, pp. 84-90; eggs, 90-94.)

COOLING, L. E. 1924. Larval stages and biology of the com- moner Australian Mosquitoes. Comm. Aust. Dept. Health Service, Melbourne, Publ. No. 8, pp. 1-40. (Paper not seen.)

DYAR, H. G. 1906. Larvae of Culicidae classified as inde- pendent organisms. Jour. N. Y. Ent. Soc., 14: 169-230. (Many keys throughout the work.)

ID. 1906. Key to the known larvae of the mosquitoes of the United States. U. S. Dept. Agr. Bur. Ent. Circ., 72: 1-6.

ID. 1928. The mosquitoes of the Americas. Carnegie hist. Wash., Pub. 387. 616 pp., 123 pis. (Various keys to larvae, no pupal keys.)

EDWARDS, F. W. 1912. Revised keys to the known larvae of African Culicinae. Bull. Ent. Res., 3:373-385. (Key to genera and species, p. 373.)

10 ENTOMOLOGICAL NEWS [Jan., '39

ID. 1912. A revision of the mosquitoes of the palaearctic region. Bull. Ent. Res., 12:263-351. (Several keys to larvae of various genera.)

ID. 1932. Culicidae. Gen. Ins., Fasc. 194, pp. 1-258. (Vari- ous larval keys to genera of the world.)

FELT, E. P. 1904. Mosquitoes or Culicidae of New York State. N. Y. Mus. Bull. 79, Ent. 22, pp. 241-400. (Larval key, p. 279.)

Fox, C. 1925. Insects and disease of man. Blakiston's Son and Co., Phila., 349 pp. (Keys to larvae from Howard, Dyar, and Knab, pp. 28, 30, and 32.)

GRUNBERG, K. 1910. [Genera of German Culicidae.] In: Brancr, Susswasserfauna Deutschlands, Heft 2A. 312 pp. Fischer, Jena. (Key to larval genera, p. 80.)

HEADLEE, T. J. 1921. Mosquitoes of New Jersey. N. J. Agr. Exp. Sta., Bull. 348, 229 pp. (Key to larvae, pp. 20-23; "eye and hand lens key" to larvae, p. 30.)

HEGNER, R., F. M. ROOT, and D. L. AUGUSTINE. 1929. Ani- mal parasitology with special reference to man and domesti- cated animals. Century Biol. Series, Century Co., N. Y., 731 pp. (Larval key to tribes, p. 482; Anopheles, p. 497; various genera, p. 520 and p. 522.)

HERMS, W. B. 1923. Medical and veterinary entomology. 2nd ed., 462 pp. Macmillan Co., N. Y. (Larval keys to Ano- pheles from Howard, Dyar and Knab, p. 126.)

HOPKINS, G. H. E. 1936. Mosquitoes of the Ethiopian region I. Larval bionomics of mosquitoes and taxonomy of culicine larvae. London. (British Museum.) 1936, 250 pp. (Various larval keys to species.)

' HOWARD, L. O., H. G. DYAR and F. KNAB. 1914-1917. The mosquitoes of North and Central America and the West Indies. 4 Vols., Carnegie Inst., Wash., D. C. (Various larval keys in Vols. 2 and 3.)

INGRAM, A. and J. W. S. MACFIE. 1917. Notes on some dis- tinctive points in the pupae of West African mosquitoes. Bull. Ent. Res., 8: 73-91. (Keys to species of pupae.)

JAMES, S. P. and W. G. LISTON. 1911. A monograph of the anopheline mosquitoes of India. Thacker, Spink & Co., Cal- cutta, 128 pp. (Key to genera and species of larvae, pp. 50-58.)

JOIIANNSEN, O. A. 1903. Culicidae. In: N. Y. St. Mus., Bull. 68, pp. 388-429. (Key to genera of larvae and pupae, pp. 391-392; species of Culex, p. 415; Aedes, p. 425; Corethra p. 393.)

(To be continued.)

1, '39] ENTOMOLOGICAL NEWS 11

A Synopsis of the Odonata of Alaska.

By LEONORA K. GLOYD, Chicago Academy of Sciences,

Chicago, Illinois.

Until recently most of the specimens of Odonata recorded from Alaska were taken by men whose task was to gather as many objects of natural history as time and facilities permitted, by collectors who were obliged to obtain as many specimens in all orders of insects as possible, or by men in other fields of activity who took time from their own work to capture a few insects for entomological friends. Naturally the Odonata, a group often difficult to catch and to preserve, would not be well represented in such collections.

The first dragonflies reported from Alaska (Hagen 1856, 1857 and 1861), representing four species, were taken between 1823 and 1839 in the vicinity of Sitka, Norton Sound and Kenai Island, by some adventurous Russian explorers who skirted the north Pacific coast-line from the Kurile Islands north of Japan to San Francisco, California. Apparently the next specimens taken (Hagen 1875, Walker 1912 and 1925) are from Fort Yukon, collected by W. H. Dall, probably during his survey of The Yukon River in 1867. In 1890 Hagen re- corded another species from Sitka but did not give the date of collection or the name of the collector. In 1894, J. A. Caden- head secured a specimen from Burrough's Bay (Walker, 1912). In 1899 members of the Harriman Alaska Expedition obtained eight species, six of which were new to the known fauna (Cur- rie 1901, 1904) ; later in the same year Rev. S. Hall Young and J. Murray Presnall collected two dragonflies from the in- terior between Mission and Forty-Mile Creeks and at Eagle City (Holland, 1900). In 1908 another species was recorded without data by Martin. Sometime prior to 1917 a species new to the fauna was taken by A. Stecker from the Kuskokwim River (Kennedy 1917 and Walker 1925). Except for a series of specimens of a previously reported species collected in 1917 by J. S. Hine at Katmai (Walker 1925), apparently no more were taken until 1933 when R. R. Sheppard, a student from

12 ENTOMOLOGICAL NEWS [Jan., '39

the University of Florida, visited Admiralty Island, securing ten species and adding three to the list (Gloyd 1938). Thus, almost a hundred years of incidental collecting yielded only a total of eighteen species. Then in the summer of 1937 Carsten Ahrens went to Alaska primarily for the purpose of collecting Odonata and, although the season was unusually cold, was suc- cessful not only in finding all but four of the previously re- ported species but also six additional ones (Ahrens 1938).

From this review of the literature it is evident that few col- lections have been recorded and that little is known about the distribution of even the most common species in the region. Many of the records are in diverse publications some of which are now difficult of access and it is hoped that the following synopsis and bibliography may facilitate the studies of others and may help to increase interest in the odonate fauna of this far north land.

References under each species are given in chronological order and those repeating earlier records are omitted in the list unless they are of taxonomic importance. As much as per- mitted by available data, the earliest and latest dates of collec- tion are also included.

LEUCORRHINIA BOREALIS Hagen. Anchorage, Gulkana (Ahrens 19386, p. 227). July 17-27.

L. HUDSONICA (Selys). Kukak Bay, Virgin Bay (Prince William Sound) (Currie 1901, pp. 221-222) ; Admiralty Island (Gloyd 1938, p. 199) ; Chitina, Gulkana, Juneau, Ketchikan (Ahrens 19386, p. 226). June 25-August 26.

L. PROXIMA Calvert. Anchorage (Ahrens 19386, p. 227). July 22.

LIBELLULA QUADRIMACULATA Linne. Fox Point (Currie 1901, p. 221, as Leptetrum quadrimaculata} ; Juneau (Ahrens 19386, p. 226). July 11-26.

SYMPETRUM DANAE (Sulzer). Admiralty Island (Gloyd 1938, pp. 198, 199) ; Juneau (Ahrens 19386, p. 226). August 5-21.

S. DECISUM (Hagen). Chitina (Ahrens 19386, p. 226). July 28.

1, '39] ENTOMOLOGICAL NEWS 13

CORDULIA SHURTLEFFI Scudder. Kukak Bay, Fox Point (Currie 1901, p. 220) ; Anchorage, Gulkana, Juneau, Palmer *(Ahrens 19386, p. 226) ; Fort Yukon.1 June 25-July 26.

SOMATOCHLORA ALBiciNCTA ( Burmeister ) . Fort Yukon (Hagen 1875, pp. 59-60, as Epitheca albicincta) ; Kodiak, Kukak Bay (Currie 1901, p. 221 ; Walker 1925, p. 172) ; Kat- mai (Walker 1925, p. 173) ; Admiralty Island (Gloyd 1938, p. 199); Anchorage, Juneau, Palmer (Ahrens 1938ft, p. 226). June 25-August 26.

S. HUDSONICA (Hagen). Fort Yukon (Walker 1925, p. 180). June 25.

S. SAHLBERGI Trybom. Kuskokwim River (Kennedy 1917, pp. 229-236, pi. 13, as walkeri n. sp. ; Walker 1925, pp. 163-167).

S. SEMICIRCULARIS (Selys). Admiralty Island (Gloyd 1938, pp. 198, 199); Juneau (Ahrens 1938ft, p. 226). July 11- August 25.

AESHNA EREMITA Scudder.2 Alaska (Martin 1908, p. 37, as clepsydra; Muttkowski 1910, p. Ill); Bethel, Kuskokwim River3 (Walker 1912, p. 126) ; Admiralty Island (Gloyd 1938, p. 199) ; Anchorage, Chitina, Gulkana, Palmer (Ahrens 1938ft, p. 226). July 19- August 19.

AE. INTERRUPTA iNTERRUPTA Walker. Admiralty Island (Gloyd 1938, pp. 198, 199) ; Ketchikan (Ahrens 1938ft, p. 226). July 8-August 19.

AE. INTERRUPTA LINEATA Walker. Chitina, Palmer (Ahrens 1938ft, p. 226). July 19-28.

AE. JUNCEA (Linne). Kenai Island, Norton Sound (Hagen 1856, pp. 369, 380; 1861, pp. 120-121) ; Kodiak, Unga Island

1 Hagen, 1875, p. 60, says "The specimen from Ft. Yukon, Alaska, quoted as C. Shurtleffi Ball, belongs to a different species," but no other reference to this specimen was found in the literature. However, in 1935, I examined 3d1 3$, collected June 25 by W. H. Ball at Fort Yukon, of C. shurtle'ffi in the U. S. National Museum.

* After the present paper was in type it was noted that Mr. Ahrens in his published list records this locality as "Matanuska Valley."

2 Currie, 1901, lists Aeslwa clepsydra (Say) (— eremite, Walker 1912, p. 119) as previously reported, but from the literature cited by him I found no definite Alaskan record.

3 Recorded as Kuskoquin River by Walker, //. cc.

14 ENTOMOLOGICAL NEWS [Jan., '39

(Shumagin Islands), Nushagak River (Currie 1901, pp. 219- 220; Walker 1912, p. 91); Bethel, Kuskokwim River3 (Walker 1912, p. 91) ; Admiralty Island (Gloyd 1938, p. 199) ; Anchorage, Gulkana, Juneau, Ketchikan, Palmer, Seward, Valdez (Ahrens 1938a, pp. 8-9, 26; 19386, p. 226). July 11- August 25.

AE. PALMATA Hagen. Kodiak, Cook Inlet (Currie 1901, pp. 219-220, as constrlcta (Say); Walker 1912, p. 163); Admir- alty* Island (Gloyd 1938, p. 199) ; Juneau, Palmer (Ahrens 19386, p. 225). July 11 -August 21. '

AE. SITCHENSIS Hagen. Sitka (Hagen 1861, pp. 119-120, type description; 18906, pp. 353-355); Burrough's Bay (Walker 1912, p. 83) ; Juneau, Palmer (Ahrens 19386, p. 225). July 19- August 6.

ANAX JUNIUS (Drury). Sitka (Hagen 1890a, p. 306); mountains between Mission and Forty-Mile Creeks, Eagle City (Holland 1900, p. 382). July 25-August 3.

CORDULEGASTER DORSALis Hagen. Sitka (Hagen 1856, pp. 367, 381 ; in Selys 1857, p. 347, type description; 1861 ; p. 116; 1875; p. 50).

AGRION RESOLUTUM Hagen. Anchorage, Gulkana, Palmer (Ahrens 19386, p. 227). July 17-27.

ENALLAGMA BOREALE Selys. Cook Inlet, Juneau, Kukak Bay (Currie 1901, pp. 218-219, as cafoerti Morse) ; Admiralty Island (Gloyd 1938, p. 199); Gulkana, Awk Lake (Juneau), Matanuska, Palmer (Ahrens 19386, p. 227). June 29-Au- gust 25.

E. CYATHIGERUM (Charpentier) . Sitka (Hagen 1856, pp. 367, 381 ; 1861, p. 87, as Agrion annexum, n. sp.) ; Admiralty Island (Gloyd 1938, p. 200); Awk Lake (Juneau), Palmer (Ahrens 19386, p. 227). July 19-August 26.

LESTES DISJUNCTUS Selys. Fox Point (Currie 1901, p. 217, as Lestes sp. ; Muttkowski 1910, p. 37); Admiralty Island (Gloyd 1938, p. 200) ; Anchorage, Gulkana, Juneau, Palmer (Ahrens 19386, p. 227). July 19-August 26.

L. DRYAS Kirby (uncatus Kirby). Chitina (Ahrens 1938, p. 227). July 28.

1. '39] ENTOMOLOGICAL NEWS 15

Hagen (1856, p. 381) mentioned Aeshna borealis Zett. (= caendea Strom) from Sitka "von Brandt in Hamburg" but I regard this record as questionable. While it is possible that this may represent A. caerulca scptentrionalis Burm., Hagen's reference is not included in the synonymy of this species by Walker (1912, p. 76) and its occurrence in Alaska has not been verified.

BIBLIOGRAPHY.

AHRENS, CARSTEN. 193&/. The commonest snakefeeder. The Alaska Sportsman, vol. 4, no. 5, pp. 8-9, 26.

ID. 19386. A List of Dragonflies (Odonata) taken in South- ern Alaska. Ent. News, vol. 49, no. 8, pp. 225-227.

CURRIE, ROLLA P. 1901. Papers from the Harriman Alaska Expedition. XXII. Entomological Results (14) : The Odonata. Proc. Washington Acad. Sci., vol. 3, pp. 217-223. Reprinted in 1904 in Harriman Alaska Expedition, vol. 8, pp. 145-153.

GLOYD, LEONORA K. 1938. Notes on some dragonflies (Odo- nata) from Admiralty Island, Alaska. Ent. News, vol. 49, no. 7, pp. 198-200.

HAGEN, HERMANN. 1856. Die Odonaten-Fauna des rus- sichen Reichs. Stettin ent. Zeitung., vol. 17, pp. 363-381.

ID. 1861. Synopsis of the Neuroptera of North America, with a list of the South American species. Smithsonian Misc. Coll., pp. xx -f 1-347.

ID. 1875. Synopsis of the Odonata of North America. Proc. Boston Soc. Nat. Hist., vol. 18, pp. 20-96.

ID. 1890a. Synopsis of the Odonata of North America, No.

2. The genus Anax. Psyche, vol. 5, Feb., pp. 303-308.

ID. 18906. Two species of Aeschna. Psyche, vol. 5, June, pp. 353-355.

HOLLAND, W. T- 1900. Alaska insects. Ent. News, vol. 11, no. 3, pp. 381-389.

KENNEDY, CLARENCE HAMILTON. 1917. A new species of Somatothlora (Odonata) with notes on the cingulata group. Canadian Entom., vol. 49, pp. 229-236, pi. 13, textfigs. 1-4, 25-47.

KIRBY, W. F. 1890. A synonymic catalogue of Neuroptera Odonata or dragonflies, with an appendix of fossil species. London, Gurney and Jackson, pp. x -j- 1-202.

MARTIN, RENE. 1906. Collections Zoologiques du Baron Edm. de Selys-Longchamps. Catalogue systematique et de- scriptif. Cordulines. Fasc. 17, pp. 1-98, pis. 1-3.

ID. 1908. Ibid. Aeschnines. Fasc. 18, pp. 1-84, pis. 1-2.

16 ENTOMOLOGICAL NEWS [Jan., '39

MUTTKOWSKI, RICHARD A. 1910. Catalogue of the Odonata of North America. Bull. Public Mus. Milwaukee, vol. 1, art. 1, pp. 1-207.

SELYS-LONGCHAMPS, Baron EDMOND DE (with H. A. Hagen). 1857. Monographic des Gomphines. Separate, pp. viii + 1-460, pis. 1-23. 1858, Mem. Soc. Sci. Liege, vol. 11, pp. 257-720.

WALKER, E. M. 1912. The North American dragonflies of the genus Aeshna. Univ. Toronto Studies, Biol. Ser. no. 11, pp. viii -f 1-213, pis. 1-28.

ID. 1925. The North American dragonflies of the genus Somatochlora. Univ. Toronto Studies, Biol. Ser. no. 26, pp. 1-202, pis. 1-35.

A New Species of Goelus Eschscholtz. (Coleoptera: Tenebrionidae).

By FRANK E. BLAISDELL, SR., Stanford Medical School and

Associate in Research, California Academy of Sciences,

San Francisco, California.

Coelus gracilis new species.

Gracilis belongs to the Globosus Group ' which also includes globosus Lee., grossus Csy. (solidus Csy., immature form) and saginatus Csy. ; also the insular species pacificus and remains of Fall. It is the smallest species of the group thus far dis- covered, some specimens scarcely exceeding in size the larger ones of Coelomorpha Csy. that occur in a similar habitat in the vicinity of Ensenada, Lower California, Mexico. All of the species belonging to the tribe Coelini burrow in sand dunes or sandy areas, under vegetation in littoral regions along the Pacific coast. Mainland species of the Globosus Group are known, from the data at hand, to occur only in the vicinity of Ensenada northward to Santa Cruz, California. It is interest- ing to discover a species as far north as Antioch, Contra Costa County, California; the author knows of no record of species belonging to the group under discussion occuring north of Santa Cruz.

1 Blaisdell, Synopsis and review of the species of Coelus. Trans. Amer. Ent. Soc., XIV, p. 318, 1919.

1, '39] ENTOMOLOCU'AL .\K\VS 17

Form oval to slightly oblong-ovate, small in size and about one-fifth longer than wide. Color nigro-piceous to ferruginous or paler due to immaturity ; antennae and palpi flavo-testaceous, legs more or less piceous. Cilia along sides of body, long, dense, erect and fulvous, with bristling hairs of the same kind on the legs ; hairs not occurring on the anterior pronotal margin and head behind the frontal suture, as in the species of the Ciliatus Group.

Head relatively moderate in size, twice as wide as length of an antenna ; epistoma relatively large and more prominent later- ally than the eyes, sides arcuately convergent anteriorly, angles broadly rounded, apical margin moderately deeply sinuate in middle two-fourths, sinus rather evenly arcuate, surface gradu- ally arising from the frontal suture, somewhat convex and more or less impressed within the apical margin lateral to the sinus, rather closely punctate especially toward the sides, punc- tures moderate in size ; frontal suture well marked, straight in middle three-fifths with lateral fifths arcuately attaining the eyes. Frons somewhat convex, broadly flattened in the central area, rather densely punctate, most so laterally and against the suture, punctures slightly larger 'and stronger than on the epi- stoma. Eyes coarsely facetted. Antennae in length equal to one-half the width of the epistoma, last four segments mod- erately compressed and somewhat gradually widened except the last ; first segment not visible from above, second and third obconical, the former slightly larger than the latter, both a little longer than wide, fourth to the seventh inclusive slightly wider at apex and briefly constricted at base, as long as wide ; last four segments forming a feebly defined club, the eighth triangular and as long as wide, ninth and tenth transverse the latter slightly wider, eleventh smaller, narrower and sub- quadrate.

Pronotum transverse, slightly more than twice as wide as long and twice as wide as the head; base less than one-half wider than apex, sinuation of the latter broad, moderately deep and rather straight in middle two-fourths, apical angles promi- nent and well rounded, margin beaded ; sides broadly arcuate and moderately convergent anteriorly, margin beaded and con- tinuously so with that of the apex; base transverse, margin thin and coriaceous, angles rather narrowly rounded ; disk mod- erately convex from side to side, almost evenly punctate, punc- tures small, well defined and separated by a distance equal to about three to six times their diameter, larger and denser in the submarginal area, the latter not widely impressed, each

18 ENTOMOLOGICAL NEWS [Jan., '39

puncture with a moderately long and more or less semi-erect hair. Sides less than moderately explanate. Surface finely alutaceous.

Elytra more or less moderately inflated posteriorly, about one-fifth longer than wide and three times as long as the pro- notum; sides subparallel and moderately arcuate, apex subo- gival ; disk strongly convex from side to side, less so in basal one-third, strongly so and arcuately declivous apically, surface moderately closely punctate, punctures separated by a distance equal to two to four times their diameter, smaller and quite simple in the central area, becoming a little larger and feebly muricate laterally and apically ; each puncture with a fine sub- recumbent, short hair which becomes coarser at the periphery.

Measurements. — (Types) Male: length 5 mm.; width 3 mm. Female : length 7 mm. ; width 4 mm.

Holotype, female, no. 4698, and allotypc, male, no. 4699, in the collection of Dr. E. C. Van Dyke, Museum of the Cali- fornia Academy of Sciences ; collected at Antioch, Contra Costa County, CALIFORNIA, April 24, 1938, by Dr. Van Dyke. Thirty-four paratypes with same data ; two are to be placed in the collection of the American Entomological Society, Phila- delphia. Largest specimen measures : Length 7 mm., width 4 mm. ; smallest : length 4.5 mm., width 2.5 mm.

The author has figured the genitalia of Coclus ciliatits Esch.,- genotype of the genus.3 Those of gracilis are of the same phylogenetic type, but show some specific differences. The aedeaga 4 of